Actually I disagree with fishing facing ventrally to the strata. According to my fish physiology classes, they orient towards the light where applicable, which is dorsal up. However, regardles of the normal fish, I can see being inverted inside a hole as being a very evolutionary manuever. Predator animals typically, due to mouth structure, cannot aim upwards. Some surface dwelling fish can, but not most reef fishes. And I can bet you that these fish don't live in darkness, but rather sit near the opening of holes in order to see the world... how else feed without other nighttime adaptations? The eyes don't look like most nighttime fishes eyes.
I think the major reason, at least as far as I can guesstimate, is that it provides an advantage in that it helps them evade predation.
Hows that for a theory?
Wade